After dreaming about an anime she used to watch as a child, Minoa Asagaya could not forget a particularly memorable scene. However, despite her best efforts, she cannot recall the name of the show. Due to this, Minoa asks for help from her fellow classmates at Sakaneko High School. Her conversation is overheard by Arisu Kamiigusa, the most popular and wealthy girl in class who is also a hardcore otaku. Yet even with her vast knowledge, Arisu does not recognize the show. After discovering that there isn't an anime club at their school, Minoa and Arisu create the Anime Research Club, as they may obtain the answer to Minoa's mystery if they gather people who share the same interest. Thus, Minoa is exposed to a bizarre new world—the world of anime!

Rikuo Uozumi has all but resigned himself to a bleak future, aimlessly working at a convenience store in Tokyo after graduating from college. His monotonous life is interrupted when the peculiar Haru Nonaka makes a lively appearance, frequently dropping by his workplace to befriend him. When Rikuo learns that an old college friend and crush, Shinako Morinome, has moved back into town, he reaches out to further their relationship. Unbeknownst to Rikuo however, Shinako is carrying painful memories from her past that were holding her back from accepting his feelings. Meanwhile, as Haru continually opens up to Rikuo, he discovers that she, much like him, is living by herself and wants to step out of her comfort zone into an uncertain future. The past lingers long in the mind, and the future remains elusive. At a crossroads along their intertwined paths, these three experience what it means to let go of their feelings of yesterday and embrace the change that tomorrow brings.