On the surface, the thought of Kyouko Hori and Izumi Miyamura getting along would be the last thing in people's minds. After all, Hori has a perfect combination of beauty and brains, while Miyamura appears meek and distant to his fellow classmates. However, a fateful meeting between the two lays both of their hidden selves bare. Even though she is popular at school, Hori has little time to socialize with her friends due to housework. On the other hand, Miyamura lives under the noses of his peers, his body bearing secret tattoos and piercings that make him look like a gentle delinquent. Having opposite personalities yet sharing odd similarities, the two quickly become friends and often spend time together in Hori's home. As they both emerge from their shells, they share with each other a side of themselves concealed from the outside world.

Throughout history, an elite group of shrine maidens known as "Toji" have saved the world from "Aratama," strange and malevolent beings bent on destroying humanity. In modern times, these warriors have been assigned to a special police squad to exterminate Aratama. The government has also set up five elite schools across the country to provide young girls the necessary sword fighting skills to eradicate these monsters and eventually join their fellow Toji in protecting the world. A student of one of those five schools, Kanami Etou is chosen to represent Minoseki Academy in a sword fighting tournament, where she meets the mysterious Hiyori Juujou. Although Kanami and Hiyori rise to the top of the tournament, their battle takes an unexpected turn, throwing the world of the Toji into chaos. Likely that the Toji are facing betrayal from within, the two are forced to flee the tournament, clashing with former comrades on the way.

Since ancient times, shrine maidens who wield swords have been exorcising "aratama," strange existences that threaten the human world. These young women who wear school uniforms and a sword are called "Toji," and they serve as an official unit within the police force as a "special religious service police squad." The government authorizes the Toji to wear swords and serve as government officials, and the government has set up five schools throughout the country for the girls to attend. In the spring, the five schools send their best Toji to compete in a tournament. As the tournament comes closer, all of the representatives rigorously train themselves in order to get stronger and improve their skills. (Source: ANN)

College student Yuuhi Amamiya's monotonous life is turned upside down with the appearance of Sir Noi Crezant, a talking lizard claiming to be a knight of justice. Crezant elaborates that the world is in peril—threatened by the mage Animus who wishes to annihilate it with his Biscuit Hammer, an enormous structure suspended in Earth's orbit. As the newly appointed Lizard Knight, Yuuhi must fight alongside other Beast Knights to protect the princess and save the planet from destruction. Unimpressed by the prospect of risking his life, Yuuhi continues his day as usual—until a golem created by Animus suddenly confronts him. Yuuhi is instantly overpowered, but his neighbor, Princess Samidare Asahina, luckily rescues him. To his surprise, Samidare's intention starkly differs from Crezant's; instead of saving the world, she aims to destroy it with her own hands. Captivated by her firm ambition, Yuuhi resolves to become stronger and help Samidare realize her goal. As the pair continues to fight against Animus' golems, Yuuhi finds himself gradually gaining an interest in life—but the chains of his past trauma may restrain him from progressing further.